User blog comment:Kogath/Monster Hunter 4 Ultimate for the Nintendo 3DS/@comment-28303251-20150226065658/@comment-17735225-20150226115326

Generally you unlock an offline version of quests that are available on that HR by reaching it.

Like you unlock village high rank Gore by reaching HR5, where you unlock him online as well.

Some monster aren't available in village at all until you reach corresponding HR online. (And most importantly their intros)