User blog comment:Kogath/Monster Hunter 4 Ultimate for the Nintendo 3DS/@comment-24995830-20150413182528/@comment-7584137-20150413183522

IMO, it is more like:

Main antagonist - Gore Magala for the Low Rank campaign, Seregios for the High Rank one. Rusted Kushala Daora is more of an unseen antagonist, just like Gogmazios and Dalamadur. That is, they are KNOWN to be a great and horrible threat, but don´t actually appear that much. Contrast this with Gore and Seregios, as they both have a much greater role, especially Gore. So...

"Main Villains" - Gore for Low Rank story, Seregios for High Rank;

"Secondary but also important Villains" - Rusted Kushala, Dalamadur and Gogmazios.

Crimson Fatalis and Cephadrome don´t really have that much of an antagonist rule IMO. At least no more than other monsters. Also, while on the subject, let us remind ourselves that this monsters aren´t actually looking to destroy our race or being "evil" (well, Gogmazios and Fatalis MAY be supernatural, but still not neccessarily "evil"). So, like the shark of Jaws and Sabor from Tarzan, they may be scary and threatening... but are still not actively evil. They cannot understand such things as good and evil. So take the "antagonist" term on the concempt of "Dangerous, not evil".